Cellulose Digestion in Ruminants

The digestive system of ruminant animals such as cow, goat, buffaloes or sheep exhibits pre‐gastric fermentation. This means the site of microbial digestion and anaerobic fermentation of cellulose occurs before digesta reaches the small intestine, the place in the digestive tract where nutrients are absorbed by the body for productive function.

Ruminants are a group of mammals, including cows, sheep, and deer, that have a unique digestive system with four compartments: the rumen, reticulum, omasum, and abomasum. They are animals which chew the cud, which is unchewed grass taken into the rumen which is returned to the mouth for chewing (regurgitation). Reticulum is responsible for regurgitation, forcing the cud back into the cow’s mouth as part of rumination.

The rumen is the largest compartment of the ruminant digestive system. It serves to store partially digested food, provides physical mixing of digesta with ruminal fluid, and is an anaerobic fermentation chamber. The ruminant and ruminal microbes possess a symbiotic relationship. First, the ruminant provides the perfect home for those microbes i.e., an anaerobic environment, perfect temperature and abundance of food. The microbes break that food down, including cellulose, which would be otherwise useless to the animal, and then use nutrients for their own growth. Eg. Fibrobacter succinogenes, Ruminococci .

Cellulose In Digestion – Herbivores, Termites, & Ruminants

Cellulose is a type of fiber that cannot be digested by the human digestive system. The presence of beta-acetal linkages in cellulose distinguishes it from starch and determines its digestion. The organisms that are capable of digesting the cellulose with their cellulase enzyme are microorganisms such as bacteria, fungi, and protozoa.

Except for a few multicellular organisms, such as termites, most higher organisms cannot digest cellulose without the help of microorganisms. In this article, we will discuss cellulose in digestion and why humans can digest starch but not cellulose.

Table of Content

  • What is Cellulose?
  • Structure of Cellulose
  • Cellulose Digestion in Ruminants
  • Cellulose Digestion in Termites
  • Cellulose Digestion in Herbivores
  • Reasons Why Humans Cannot Digest Cellulose
  • Importance of cellulose in digestion

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What is Cellulose?

Cellulose is a natural, linear polymer made of glucose units. It is a major component of the cell walls of most plants and is also found in some algae and bacteria. Cellulose comprises around 33% of all vegetable matter and is the most abundant of all naturally occurring organic polymer on Earth. The glucose units in cellulose are connected in a unique way (beta‐linkages) to make cellulose very resistant to breakdown. Mammals lack the enzymes required to break these beta‐linkages, required to release the glucose. But, bacteria, fungi, and protozoa that reside in the rumen of ruminant animals do possess the enzymes necessary to break those glycosidic bonds. Herbivores also possess the same microbes that can break down cellulose, but they are located in a different part of the gut known as the cecum. The relationship between microbes and herbivores is called a symbiotic relationship....

Structure of Cellulose

Cellulose is an organic compound derived from D-glucose units and having the formula (C6H10O5)n. It is a polysaccharide comprising of a linear chain of several hundred to many thousands of β(1→4) linked glycosidic bonds. This linkage motif differs from that of α(1→4)-glycosidic bonds present in starch and glycogen. Glucan chains of cellulose aggregate via hydrogen bonds and van der Waals forces to form a long thread-like crystalline structure called a cellulose microfibril....

Cellulose Digestion in Ruminants

The digestive system of ruminant animals such as cow, goat, buffaloes or sheep exhibits pre‐gastric fermentation. This means the site of microbial digestion and anaerobic fermentation of cellulose occurs before digesta reaches the small intestine, the place in the digestive tract where nutrients are absorbed by the body for productive function....

Cellulose Digestion in Termites

Animals such as termites consume wood and dry leaves which also contain cellulose. The digestion of cellulose in termites is also done by the enzyme cellulase produced by protozoans which live symbiotically with the termites. The byproducts of the breakdown of cellulose could be acetic acid or glucose....

Cellulose Digestion in Herbivores

Plants have cellulose in their cell walls, which is consumed by herbivores. Numerous herbivores use the ability of some bacterial groups to ferment plant cell walls in order to extract energy from them. After that, the body absorbs it and uses it as nourishment. Herbivores can digest cellulose by monogastric digestion by using the symbiotic gut bacteria. The efficiency of herbivores in obtaining energy from the digestion of cellulose is lower than that of ruminants. Here, microbial fermentation is used to break down cellulose....

Reasons Why Humans Cannot Digest Cellulose

Humans lack the ability to digest cellulose for several reasons:...

Importance of Cellulose in Digestion

Although cellulose itself is not directly digestible by humans, its presence in the diet has important roles for digestive health:...

FAQs -Cellulose in Digestion

How does Cellulose Help in Digestion?...

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